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The study of semiotics underwent a gradual but radical paradigm shift during the past century, from a glottocentric enterprise to one that encompasses the whole terrestrial biosphere. In this collection of seventeen essays, Tom Sebeok, one of the seminal thinkers in the field, shows how this progression took place. His wide-ranging discussion of the evolution of the field covers many facets, including discussions of bio-semiotics, semiotics as a bridge between the humanities and natural sciences, semiosis, non-verbal communication, cat and horse behaviour, the semiotic self, women in semiotics. This thorough account will appeal to season scholars and neophytes alike.